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Breeding

How Gentoo Penguins Swim So Fast Underwater: The Physics Explained

Gentoo penguins “fly” underwater with bending hydrofoil wings. Both upstroke and downstroke produce thrust, while wing angle, buoyancy and drag set their speed.

By Animalso Team 5 min read
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Gentoo penguins move quickly underwater by flying through the water with their short, flattened wings. Each wing works as a hydrofoil: as it sweeps through the water, it produces a force with a forward component. Unlike many wing-propelled divers, gentoos can gain forward acceleration on both the downstroke and the upstroke. Wing bending, feather control, buoyancy and drag determine how much useful thrust each stroke produces.

They use their wings as hydrofoils

A gentoo’s flipper is a stiff, streamlined lifting surface rather than a paddle used mainly to push water backward. Its thick aerofoil section is supported by the arm and hand bones; the 2021 study reported a maximum wing thickness of 17.5% of the chord. As the wing moves through water, pressure differences around the aerofoil generate hydrodynamic force. By orienting that force appropriately, the penguin obtains forward thrust.

Penguin feet and legs can help with steering, braking and maneuvering, but the available findings identify wing beats as the principal source of forward propulsion.

Why both halves of the stroke can push the bird forward

Downstroke

On the downstroke, the wing’s path and angle of attack create a force that has a forward component. The penguin adjusts the wing’s orientation rather than simply driving it straight down, allowing part of the hydrodynamic force to overcome drag.

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Upstroke

Gentoo penguins also produce forward acceleration while the wing rises. Three-dimensional video of three birds swimming horizontally at Nagasaki Penguin Aquarium showed forward acceleration during each half of the stroke cycle. The upstroke’s contribution is unusual among wing-propelled diving animals and is associated with large supracoracoideus (pectoral) muscles that power the wing’s return stroke.

This two-sided propulsion lets a gentoo maintain thrust through the complete cycle instead of treating the upstroke only as recovery.

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Wing bending makes the stroke more effective

A flipper does not remain a perfectly flat plate. It bends during the beat, changing the local angle of attack—especially on the upstroke. That changes how water flows over the wing and how much of the resulting force points forward.

In the 2021 work, the observed bending motion was reconstructed and compared with a flattened version of the same wing. Their quasi-steady hydrodynamic model, informed by water-tunnel measurements of a 3D-printed rigid wing, estimated 1.8 times the propulsive efficiency for the reconstructed bending wing. The figure is a comparison between two modeled wing shapes, not a direct measurement of whole-animal efficiency.

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The same analysis found higher stroke-averaged thrust for the bending shape. In practical terms, bending appears to help the penguin keep a useful angle of attack as the wing reverses direction.

Angle of attack and feathering control thrust

Angle of attack is the angle at which the wing meets the oncoming water. Too little angle produces little force; too much can increase separation and wasted drag. Active feathering—rotating or pitching the wing during the stroke—helps keep the force directed usefully as the wing’s speed and path change.

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A separate computational study, described by the American Institute of Physics, proposed an “angle of thrust” concept linking thrust to angle of attack and to the wing’s angle relative to the forward direction. Its author, Prasert Prapamonthon, said: “Thrust is primarily determined by the angle of attack and the relative angle of the wings to the forward direction.” That statement describes the study’s computational model; it should not be treated as a universal law established for every penguin or swimming condition. The work also anticipated extending the model to a more realistic three-dimensional penguin.

How buoyancy and drag set the direction of force

Water is more than 800 times denser than air, so a flipper moving at speed experiences substantial hydrodynamic loads. The same density also makes buoyancy a major part of underwater movement.

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Horizontal swimming

When a gentoo swims level, its thrust must do two jobs at once: the forward component balances drag, while a downward component can counter the bird’s tendency to rise. The penguin therefore cannot aim every bit of wing force directly forward. Wing pitch, bending and body angle determine the balance between forward motion and vertical control.

Why streamlining matters

Drag rises with speed and depends on body shape, posture and the flow around the flippers. A streamlined body reduces the force that the wings must overcome, while controlled wing motion supplies thrust without creating unnecessary sideways or vertical losses.

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How fast can gentoo penguins swim?

The best-supported figures in the cited studies describe average or cruising movement, not a verified maximum sprint. One source gives an average gentoo foraging speed of 2.3 m/s (about 8.3 km/h). A free-ranging comparison covering seven penguin species reported cruising speeds from 1.8 to 2.3 m/s (about 6.5 to 8.3 km/h). That range spans species and study conditions, so it is not a gentoo-specific top-speed measurement.

A comparative swimming model found that relationships among body mass, cruising speed and stroke frequency were consistent with minimizing metabolic transport cost. Such an energetically economical cruising speed is a different quantity from a brief escape burst. The often-repeated 36 km/h figure should not be presented as a verified maximum on the evidence summarized here.

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What is being measured Reported value or result What it means
Gentoo foraging speed 2.3 m/s average An average field value, not a maximum sprint
Comparative penguin cruising 1.8–2.3 m/s across seven species A cross-species range under the study’s conditions
Wing-shape efficiency model 1.8× for bending versus flattened wing A modeled comparison, not whole-animal efficiency

What the evidence actually shows

Evidence type What was done What can be concluded
Direct observation Multiple underwater cameras recorded three gentoo penguins swimming forward and horizontally at Nagasaki Penguin Aquarium on 7 March 2018 and 4 September 2019. The birds accelerated during both upstroke and downstroke, and their three-dimensional wing motion could be reconstructed.
Hydrodynamic calculation The analysis combined reconstructed motion, water-tunnel tests of a 3D-printed rigid wing and a quasi-steady model. The bending motion produced greater modeled thrust and an estimated 1.8-fold efficiency advantage over a flattened comparison wing.
Separate computational model A model examined flapping, feathering and wing angles. It offers a proposed explanation for how wing orientation controls thrust; its results remain model-specific.

The aquarium observations establish kinematics under those conditions. They do not by themselves provide a wild gentoo’s maximum speed or a direct measurement of metabolic efficiency. Likewise, modeled forces depend on the reconstructed motion, wing geometry and quasi-steady assumptions.

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Putting the swimming strategy together

  1. Generate lift and thrust: the flippers act as hydrofoils moving through dense water.
  2. Use the whole stroke: both downstroke and upstroke can accelerate the bird forward.
  3. Adjust the wing: bending and active feathering change angle of attack and force direction.
  4. Balance the environment: forward thrust offsets drag while vertical force helps manage buoyancy.
  5. Choose an economical pace: cruising and foraging speeds reflect a balance between transport cost and the thrust required, not necessarily the bird’s brief maximum.

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